The Aquatic Microbial Ecology Research Group at the University of Tennessee



Steven W. Wilhelm, Ph.D. (Western Ontario)

Professor of Microbiology and benevolent dictator

Professor Wilhelm is the laboratory PI and responsible for herding cats, chaos control, wine selection and middle relief.

Steven Wilhelm is the Kenneth and Blaire Mossman Professor of the Department of Microbiology.  Professor Wilhelm is a Fellow of the American Academy of Microbiology as well as a Sustaining Fellow of the Association for the Sciences of Limnology and Oceanography. In 2018 he was also named a James R. Cox Professor at the University of Tennessee, a position he held through 2021. Recently he was named the winner (along with Curtis Suttle) of the John H Martin Award from ASLO for their initial description of  the viral shunt.

Gary LeCleir, Ph.D. (Georgia)  

Research Assistant Professor, REU Director

Dr LeCleir came to Knoxville in 2006 after completing his PhD at the University of Georgia.  He is our in house statistical guru, field deployment leader and reigning karaoke champion.  Gary is also the in house expert on  all things Boston Red Sox and manages the oUTsiders softball team.

Kristin Yoshimura, Ph.D. (Delaware)  

Postdoctoral Associate

Kristen joined the lab mid-quarantine (July 2020) and will be working on our Department of Energy OBER-funded project to examine viruses of protists and how the effects of climate variation can be placed into descriptive models of system function.

Robbie Martin, Ph.D. (Tennessee)     


Postdoctoral Associate


Robbie joined the lab in late 2012 as a PhD student and has been a virtual Jack-of-all-trades.  After completing his PhD he accepted our offer to stay on to continue to work on our toxic cyanobacterial projects.  Presently Robbie is setting running chemostat studies of Microcystis gene regulation and physiology that will feed into modeling studies by collaborators at  the Technical University of Berlin.

Emily Chase, Ph.D. (Aix-Marseille Université)       

Postdoctoral Associate

Emily joined the lab in early 2022 after completing her doctoral work in France as well as a MSc and BSc at Acadia (Canada).  She has joined the group to work on our AaV genetics team, seeking to develop tools to make the genome of this virus tractable.


Naomi Gilbert,  B.Sc. (James Madison)       

Doctoral Candidate, Microbiology

Naomi came to the lab in the fall of 2017 from James Madison University (and former AMERG member Morgan Steffen's lab).  Naomi's dissertation research will see her collecting samples in the southern ocean as part of a research collaboration with scientists at CSIRO (Hobart) and Bigelow.  Her focus will be to resolve virus-host interactions from transcriptional data sets as well as to characterize the interplay between bacteria and phytoplankton with respect to trace metal cycling in marine surface waters.

Brittany Zepernick,  B.Sc. (Bowling Green)       

Doctoral Candidate, Microbiology

Brittany joined the lab in fall 2018 as part of our harmful algal blooms team.  She will be working on our collaborations with researchers in Ohio, Michigan and NC to examine factors that drive Microcystis success and ecological outcomes in large lakes.  Along with being a NSF-GRFP winner, she holds a Tennessee Fellowship for Graduate Excellence.  Most recently she has jumped into the world of freshwater diatoms to try and understand why they cannot compete with Microcystis.

Liz Denison,  M.Sc. (UNC Wilmington)       

Doctoral Candidate, Microbiology

Liz joined the lab in fall 2019 after completing a MSc project looking at cetacean microbiomes.  Currently she is working with our collaborators at ORNL and Duke on a project to examine virus activity in Sphagnum bogs that are part of the DOE Project Spruce initiative.   Along with examining the effects of temperature on virus infection, Liz is becoming our in-house expert on all-things mitovirus. Currently she holds a UT GATE Fellowship for her work.


Gwendolyn Stark,  B.Sc. (SUNY ESF)       

Doctoral Candidate, Microbiology

Gwen came to the lab in fall 2019 with an interest in toxic cyanobacteria and fresh waters.   Presently she is working with Robbie Martin on the cold temperature phenotype in Microcystis as well as resolving strain variability that we see between different Microcystis isolates.

Alex Truchon,  M.Sc. (U Mass Amherst)       

Doctoral Candidate, Microbiology

Alex joined the lab in the fall of 2020 where he is working as part of the Aureococcus team.  Along with working on resequencing the virus and some hosts, he will be continuing our efforts to make the AaV virus-host system genetically tractable. Alex was awarded a Tennessee Fellowship for Graduate Excellence when he joined UT.

Barbara Klein,  B.Sc. (U West Florida)       

Doctoral Candidate, Microbiology

Barb is a former UTK REU student that has returned to Knoxville to continue her research.  Out of the gate she will be working with Microcystis in chemostats and trying to resolve how viruses are involved in horizontal gene transfer.



Katelyn Hougton,  M.Sc. (U West Florida)       

Doctoral Candidate, Microbiology

Katelyn joined the lab in late 2021 to work with us on our projects examining virus - host interactions.  She will be using single-celled approaches to transcriptomics and samples from the Sargasso Sea to tease apart "who infects who".



David Niknejad    

Undergraduate Assistant, Microbiology

David is a UTK Microbiology major who will be working with Brittany on her efforts to examine competition between Microcystis and diatoms. 

Laura Smith, M.Sc. (Delaware)

Research Associate

With a BSc from The Ohio State University and a MSc from Delaware, Laura has joined the lab in early 2021 to help our team working on all thing Microcystis. Along with maintaining culture stocks she will be helping to get our various chemostats up and running (and keep them that way) for experiments ranging from long term evolution influences to the effects of episodic events.




Graduate student alumni


Helena Pound

Microbial Community Dynamics of a Microcystis Bloom (Ph.D. 2021)

Eric Gann

Physiological and ecological characterization of the Aureococcus anophagefferens virus host system (Ph.D. 2020)

Samantha Coy

Tool development for model studies on the interactions between a eukaryotic algae and a giant virus (Ph.D. 2019)

Robbie M. Martin

Ecological constraints of toxic cyanobacterial blooms (Ph.D. 2018)

Lauren E. Krausfeldt

Molecular characterization of factors constraining the success and toxicity of Microcystis blooms (Ph.D. 2018)

P. Jackson Gainer

Microbial interactions in the North Pacific Ocean (Ph. D. 2018)

Joshua M. A. Stough

Prediction of host-microbe interactions from community high-throughput sequencing data (Ph.D. 2017)

Mohammad Moniruzzaman

Molecular and ecological aspects of interactions between Aureococcus anophagefferens and its giant virus (Ph.D. 2016)

Morgan Steffen 

Systems biology of Microcystis blooms (Ph.D. 2014)

Shafer Belisle 

Urea as a nutrient source for Lake Erie blooms (M.Sc. 2014)

Tiana Pimentel 

Environmental constraints on cyanomyophage in the Pacific Ocean (M.Sc. 2013)

Claire Campbell

The effects of nutrient limitation and cyanophage on heterotrophic microbial diversity (M.Sc. 2012)

Audrey Matteson

Quantification and ecological perspectives on cyanophage and aquatic viruses (Ph.D. 2011)

Matthew Saxton   

Constraints on primary production in Lake Erie (Ph.D. 2011)

Star Loar  

Seasonal Variation in Lake Erie picoplankton (M.Sc. 2009)

Janet Rowe 

Ecological aspects of viruses in marine systems (Ph.D. 2008)

Johanna Rinta-Kanto

Biogeography and genetic diversity of toxin producing cyanobacteria in a Laurentian Great Lake (PhD. 2006)

Leo Poorvin

The role of viruses in Fe cycling in the World's oceans (Ph.D. 2005)

Julie L. Higgins

Virus dynamics in high nutrient - low chlorophyll marine surface waters (M.Sc. 2005)

Cecile E. Mioni

Using a bioluminescent bacterial bioreporter to assess iron bioavailability in the ocean (Ph.D. 2004)

Melanie Eldridge

The effects of Fe on plankton in HNLC regions of the world's oceans (Ph.D. 2004)

Amanda L. Dean

The dynamics, distribution, and activity of viruses in Lake Erie (M.Sc. 2004)

Shannon Pedigo-Eftland

The effects of iron on growth and physiology of the cyanobacterium Microcystis aeruginosa (M.Sc. 2004)

Johanna Rinta-Kanto


The effects of viral size class enrichments on microbial communities in marine systems   (M.Sc. 2001)



updated 01/27/2022